What is a Cetacean?
Lesson 1 · Mammals that chose the ocean
Articles and studies
- A diminutive new basilosaurid whale reveals the trajectory of the cetacean life histories during the Eocene (2023) – Communications Biology
- Basilosaurus, the “King of Lizards” (2020) – Whale Scientists
- Hippos and Whales: Unlikely Cousins (2017) – Royal Ontario Museum
- Science Word of the Day: Involucrum (2015) – National Geographic
- Pakicetus (2022) – EBSCO Research Starters
- From Land to Water: the Origin of Whales, Dolphins, and Porpoises (2009) – Evolution: Education and Outreach
- New Protocetid Whale from the Middle Eocene of Pakistan: Birth on Land, Precocial Development, and Sexual Dimorphism (2009) – PLoS ONE
- Whales originated from aquatic artiodactyls in the Eocene epoch of India (2007) – Nature
- The Origin of Whales and the Power of Independent Evidence (2000) – NCSE
- Phylogenetic relationships among cetartiodactyls based on insertions of short and long interspersed elements: Hippopotamuses are the closest extant relatives of whales (1999) – PNAS
- The evolution of whales – Understanding Evolution, UC Berkeley
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- Spying on Whales (2018) – Nick Pyenson
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Lesson 2 · Built for water
Articles and studies
- A Whale of a Tail: Unraveling the evolutionary secrets of how whales and dolphins adapted their backbones for aquatic life (2024) – Harvard University
- The Biological Productivity of the Ocean (2012) – Nature Education
- Species review of Cuvier’s beaked whale, Ziphius cavirostris (2011) – International Whaling Commission
- 5.1 Properties of Water – Introduction to Oceanography
- Properties of Water – ResearchGate
- Whales – Smithsonian Ocean
- The loudest voice in the animal kingdom – BBC Earth
- Do whales and dolphins have hair? – Whale and Dolphin Conservation
- Mammal – ScienceDirect Topics
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
- Cetacean Societies: Field Studies of Dolphins and Whales (2000) – Mann, Connor, Tyack & Whitehead (eds.)
Lesson 3 · Two great lineages
Articles and studies
- Cetaceans Have Complex Brains for Complex Cognition (2007) – PLoS Biology
- Intro to Odontocetes & Mysticetes (2017) – Ocean Conservation Research
- Vaquita – NOAA Fisheries
- Intelligence, Cognition and Emotion in Cetaceans – The Whale Sanctuary Project
- Are Whales Sentient? The Scientific Evidence (2026) – Biology Insights
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- Cetacean Societies: Field Studies of Dolphins and Whales (2000) – Mann, Connor, Tyack & Whitehead (eds.)
Lesson 4 · Underwater living
Articles and studies
- Nursing Behavior in Sperm Whales (Physeter macrocephalus) (2023) – Animal Behavior and Cognition
- Fight or flight: antipredator strategies of baleen whales (2008) – Mammal Review
- Cetacean sleep: An unusual form of mammalian sleep (2008) – Neuroscience and Biobehavioral Reviews
- Lactation in Whales and Dolphins: Evidence of Divergence Between Baleen- and Toothed-Species (1997) – Journal of Mammary Gland Biology and Neoplasia
- [TITLE MISSING] – PubMed Central
- Blue Whale Life Cycle: Birth, Growth, and Lifespan (2026) – Science Insights
- Intelligence, Cognition and Emotion in Cetaceans – The Whale Sanctuary Project
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
The Great Filter Feeders
Lesson 1 · The blue whale
Articles and studies
- Colossal ears? How baleen whales hear low-frequency sound (2025) – Hearing Research
- Blue whales (2020) – Current Biology
- Insight into the kinematics of blue whale surface foraging through drone observations and prey data (2020) – PeerJ
- Balaenoptera musculus (Blue Whale) (2018) – IUCN Red List
- A pulsed-air model of blue whale B call vocalizations (2017) – Scientific Reports
- Worldwide decline in tonal frequencies of blue whale songs (2009) – Endangered Species Research
- Studying How Cetaceans Use Sound to Explore Their Environment (1997) – Perspectives in Ethology
- Blue Whale – NOAA Fisheries
- Blue whale – International Whaling Commission
Lesson 2 · The humpback whale
Articles and studies
- Southern Ocean humpback whales are shifting to an earlier return migration (2025) – Scientific Reports
- Study finds humpback whales only ones capable of bubble-net feeding (2025) – University of Hawaiʻi
- Whales always had voices. Roger Payne helped the world hear them (2023) – Greenpeace
- Why Send Whale Song Into Space? (2017) – Literary Hub
- Humpback whales interfering when mammal-eating killer whales attack other species: Mobbing behavior and interspecific altruism? (2017) – Marine Mammal Science
- Cultural revolution in whale songs (2000) – Nature
- How do the ventral grooves of rorquals function? – Baleines en direct
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Lesson 3 · Right whales
Articles and studies
- Extreme longevity may be the rule not the exception in Balaenid whales (2024) – Science Advances
- Decadal decline in maternal body condition of a Southern Ocean capital breeder (2023) – Scientific Reports
- Saving the North Atlantic right whale in a changing ocean: Gauging scientific and law and policy responses (2020) – Ocean & Coastal Management
- Remote Acoustic Monitoring of North Atlantic Right Whales (Eubalaena glacialis) Reveals Seasonal and Diel Variations in Acoustic Behavior (2014) – PLoS ONE
- Swim Speed, Behavior, and Movement of North Atlantic Right Whales (Eubalaena glacialis) in Coastal Waters of Northeastern Florida, USA (2013) – PLoS ONE
- Dangerous dining: surface foraging of North Atlantic right whales increases risk of vessel collisions (2011) – Biology Letters
- Observations of a Female North Atlantic Right Whale in Simultaneous Copulation with Two Males: Supporting Evidence for Sperm Competition (2005) – Aquatic Mammals
- 10 Things You Should Know About North Atlantic Right Whales – NOAA Fisheries
- Protecting North Atlantic right whales from collisions with vessels in the Gulf of St. Lawrence – Transport Canada
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 4 · Gray whales
Articles and studies
- An extremely rare gray whale was spotted in the Atlantic. Scientists say it’s only the beginning. (2024) – National Geographic
- Fish Feeding and Rapid Foraging Behavior Switching by Gray Whales (Eschrichtius robustus) in California (2024) – Aquatic Mammals
- Effects of vessel traffic and ocean noise on gray whale stress hormones (2022) – Scientific Reports
- Natal origin of Namibian grey whale implies new distance record for in-water migration (2021) – Biology Letters
- Migrating eastern North Pacific gray whale call and blow rates estimated from acoustic recordings, infrared camera video, and visual sightings (2019) – PLoS ONE
- Tongue and Hyoid Musculature and Functional Morphology of a Neonate Gray Whale (Cetacea, Mysticeti, Eschrichtius robustus) (2015) – The Anatomical Record
- What Happened to Gray Whales during the Pleistocene? The Ecological Impact of Sea-Level Change on Benthic Feeding Areas in the North Pacific Ocean (2011) – PLoS ONE
- Gray whales may increase feeding opportunities for avian benthivores (2008) – Marine Ecology Progress Series
- Fight or flight: antipredator strategies of baleen whales (2008) – Mammal Review
- Side-scan sonar estimates of the utilization of gray whale feeding grounds along Vancouver Island, Canada (1986) – R. Kvitek & J. Oliver
Lesson 5 · The bowhead whale
Articles and studies
- DNA repair and anti-cancer mechanisms in the long-lived bowhead whale (2023) – bioRxiv (preprint)
- Accuracy of the Aspartic Acid Racemization Technique in Age Estimation of Mammals and the Influence of Body Temperature (2018) – NAMMCO Scientific Publications
- Oral cavity hydrodynamics and drag production in Balaenid whale suspension feeding (2017) – Semantic Scholar
- Insights into the Evolution of Longevity from the Bowhead Whale Genome (2015) – Cell Reports
- The transcriptome of the bowhead whale Balaena mysticetus reveals adaptations of the longest-lived mammal (2014) – Aging
- Hydrodynamic and Sensory Factors Governing Response of Copepods to Simulated Predation by Balaenid Whales (2012) – Semantic Scholar
- Models of hydrodynamic flow in the bowhead whale filter feeding apparatus (2004) – Journal of Experimental Biology
- Observations on the Ice-Breaking and Ice Navigation Behavior of Migrating Bowhead Whales (Balaena mysticetus) near Point Barrow, Alaska, Spring 1985 (1988) – JSTOR
Lesson 6 · Filter feeding
Articles and studies
- How Baleen Whales Feed: The Biomechanics of Engulfment and Filtration (2017) – Annual Review of Marine Science
- Metabolic Expenditures of Lunge Feeding Rorquals Across Scale: Implications for the Evolution of Filter Feeding and the Limits to Maximum Body Size (2012) – PLoS ONE
- Mechanics, hydrodynamics and energetics of blue whale lunge feeding: efficiency dependence on krill density (2011) – Journal of Experimental Biology
Toothed Whales
Lesson 1 · Dolphins
Articles and studies
- Some Dolphins Use Tools to Hunt, Revealing Insights Into Cultural Transmission (2025) – Georgetown University
- Episodic-like memory in common bottlenose dolphins (2022) – Current Biology
- Evidence that bottlenose dolphins can communicate with vocal signals to solve a cooperative task (2021) – Royal Society Open Science
- Fine-scale sociality reveals female–male affiliations and absence of male alliances in bottlenose dolphins (Tursiops truncatus) in the Shannon Estuary, Ireland (2019) – Marine Mammal Science
- Decades-long social memory in bottlenose dolphins (2013) – Proceedings of the Royal Society B
- Bottlenose dolphins can use learned vocal labels to address each other (2013) – PNAS
- Cognitive skills in bottlenose dolphin communication (2013) – Trends in Cognitive Sciences
- Dolphin cognition (2004) – Current Biology
Books
- The Animal Ethics Reader (2003) – Armstrong & Botzler (eds.)
Lesson 2 · Sperm whale
Articles and studies
- Sperm whale clans and human societies (2024) – Royal Society Open Science
- Contextual and combinatorial structure in sperm whale vocalisations (2024) – Nature Communications
- Sperm Whale Phonetic Alphabet Proposed for the First Time (2024) – Project CETI
- Deep Machine Learning Techniques for the Detection and Classification of Sperm Whale Bioacoustics (2019) – Scientific Reports
- Diving and Feeding Behaviour of Sperm Whales (Physeter macrocephalus) in the Northwestern Mediterranean Sea (2004) – Aquatic Mammals
- The face that sank the Essex: potential function of the spermaceti organ in aggression (2002) – Journal of Experimental Biology
- Research – Project CETI
Lesson 3 · Beaked whales
Articles and studies
- Navigating the unknown: assessing anthropogenic threats to beaked whales, family Ziphiidae (2024) – Royal Society Open Science
- Behavioural Ecology Traits of Elusive Deep-Diver Whales Unravel a Complex Social Structure Influenced by Female Philopatry and Defence Polygyny (2022) – Frontiers in Marine Science
- Fear of Killer Whales Drives Extreme Synchrony in Deep Diving Beaked Whales (2020) – Scientific Reports
- Diving behaviour of Cuvier’s beaked whales (Ziphius cavirostris) off Cape Hatteras, North Carolina (2019) – Royal Society Open Science
- Advances in research on the impacts of anti-submarine sonar on beaked whales (2019) – Proceedings of the Royal Society B
- Microplastic and macroplastic ingestion by a deep diving, oceanic cetacean: The True’s beaked whale Mesoplodon mirus (2015) – Environmental Pollution
- No mass strandings since sonar ban (2013) – Nature
- Beaked whales respond to simulated and actual navy sonar (2011) – PLoS ONE
- Correlating military sonar use with beaked whale mass strandings: what do the historical data show? (2009) – Aquatic Mammals
- Extreme diving of beaked whales (2006) – Journal of Experimental Biology
- Elements of beaked whale anatomy and diving physiology and some hypothetical causes of sonar-related stranding (2006) – Journal of Cetacean Research and Management
- Understanding the impacts of anthropogenic sound on beaked whales (2005) – Journal of Cetacean Research and Management
Lesson 4 · Pilot whales
Articles and studies
- Life History Parameters, Sociobiology, and Reproductive Strategies of Pilot Whales (2023) – Sex in Cetaceans (book chapter)
- Ritual Slaughter: The Tradition of Pilot Whale Hunting on the Faroe Islands (2021) – Frontiers in Veterinary Science
- Underlying environmental factors and geomagnetic fields linked to 45 pilot whales Globicephala macrorhynchus stranding in Modung white beach, Indonesian coast on 19th February 2021 (2021) – bioRxiv (preprint)
- Vocal repertoire of long-finned pilot whales (Globicephala melas) in northern Norway (2017) – Journal of the Acoustical Society of America
- The Mass Stranding Event of Long-finned Pilot Whales at Golden Bay in February 2017 (2017) – Kokako
- Mobbing-like behavior by pilot whales towards killer whales: a response to resource competition or perceived predation risk? (2015) – Acta Ethologica
- Pilot Whales Attracted to Killer Whale Sounds: Acoustically-Mediated Interspecific Interactions in Cetaceans (2012) – PLoS ONE
- Satellite tracking of long-finned pilot whales (Globicephala melas) following stranding and release in Tasmania, Australia (2012) – Wildlife Research
- The role of synchronized swimming as affiliative and anti-predatory behavior in long-finned pilot whales (2012) – Behavioural Processes
- Synchronous breathing by pilot whales (2012) – Marine Mammal Science
Lessons 5–6 · Porpoises & River dolphins
Articles and studies
- Estimating the abundance of the critically endangered Baltic Proper harbour porpoise (Phocoena phocoena) population using passive acoustic monitoring (2022) – Ecology and Evolution
- A case of playful interaction between Bolivian river dolphins with a Beni anaconda (2022) – JSTOR
- River Dolphins Spotted in Rare Playful Interaction With a Beni Anaconda (2022) – Smithsonian Magazine
- Polluted porpoises: Generational transfer of organic contaminants in harbour porpoises from the southern North Sea (2021) – Science of the Total Environment
- Interacting effects of vessel noise and shallow river depth elevate metabolic stress in Ganges river dolphins (2019) – Scientific Reports
- Porpoises the World Over: Diversity in Behavior and Ecology (2019) – Ethology and Behavioral Ecology of Odontocetes (book chapter)
- High field metabolic rates of wild harbour porpoises (2018) – Journal of Experimental Biology
- Morphological convergence in ‘river dolphin’ skulls (2017) – PeerJ
- Ultra-High Foraging Rates of Harbor Porpoises Make Them Vulnerable to Anthropogenic Disturbance (2016) – Current Biology
- The Neocortex of Indian River Dolphins (Genus Platanista): Comparative, Qualitative and Quantitative Analysis (2016) – Brain, Behavior and Evolution
- Do Porpoises Choose Their Associates? A New Method for Analyzing Social Relationships among Cetaceans (2011) – PLoS ONE
- Coexistence of Fisheries with River Dolphin Conservation (2010) – Conservation Biology
- Bye Baiji? (2007) – Current Biology
- River Dolphins Initiative – WWF
- Myths and Legends of the Amazon’s Pink River Dolphin – South America Vacations
Lesson 7 · Echolocation
Articles and studies
- For Dolphins, Echolocation May Be More Like ‘Touching’ Than ‘Seeing’ (2025) – Scientific American
- Adaptive echolocation behavior of bats and toothed whales in dynamic soundscapes (2023) – Journal of Experimental Biology
- Echolocating toothed whales use ultra-fast echo-kinetic responses to track evasive prey (2021) – eLife
- A bigger nose, a bigger bang: size matters for echolocating toothed whales (2018) – Aarhus University
- Clicking in Shallow Rivers: Short-Range Echolocation of Irrawaddy and Ganges River Dolphins in a Shallow, Acoustically Complex Habitat (2013) – PLoS ONE
- The Acoustic World of Harbor Porpoises – American Scientist
- Possible limitations of dolphin echolocation: a simulation study based on a cross-modal matching experiment – no link
Lesson 8 · Orca
Articles and studies
- Why these orcas are wearing salmon as hats (again) (2024) – National Geographic
- Ancestry testing of “Old Tom,” a killer whale central to mutualistic interactions with human whalers (2023) – Journal of Heredity
- These orcas control the waves to hunt. It’s spine-tingling to watch. (2023) – National Geographic
- Independent acoustic variation of the higher- and lower-frequency components of biphonic calls can facilitate call recognition and social affiliation in killer whales (2020) – PLoS ONE
- Genome-culture coevolution promotes rapid divergence of killer whale ecotypes (2016) – Nature Communications
- The Cooperation of Humans and Killer Whales (Orcinus orca): The Application of a Simple Fuzzy Rule-Based Model to a Historical System (2015) – Academia.edu
- Killer whale ecotypes: Is there a global model? (2013) – Biological Reviews
- Cultural traditions and the evolution of reproductive isolation: ecological speciation in killer whales? (2012) – Biological Journal of the Linnean Society
- Antarctic Peninsula killer whales (Orcinus orca) hunt seals and a penguin on floating ice (2012) – Marine Mammal Science
- Cross-cultural and cross-ecotype production of a killer whale ‘excitement’ call suggests universality (2011) – Naturwissenschaften
- Killer whales (Orcinus orca) feeding on schooling herring (Clupea harengus) using underwater tail-slaps: kinematic analyses of field observations (2000) – Journal of Experimental Biology
- Development of intentional stranding hunting techniques in killer whale (Orcinus orca) calves at Crozet Archipelago (1995) – Canadian Journal of Zoology
- Surface and underwater observations of cooperatively feeding killer whales in northern Norway (1993) – Canadian Journal of Zoology
- Intentional stranding apprenticeship and social play in killer whales (Orcinus orca) (1991) – Canadian Journal of Zoology
- Acoustic behaviour of resident killer whales (Orcinus orca) off Vancouver Island, British Columbia (1989) – Canadian Journal of Zoology
Books
- Endangered Orcas: The Story of the Southern Residents (2019) – Monika Wieland Shields
Lives in the Ocean
Lesson 1 · Migration
Articles and studies
- How Marine Mammals Navigate – A Review (2026) – University of British Columbia, Working Paper
- Migrating baleen whales transport high-latitude nutrients to tropical and subtropical ecosystems (2025) – Nature Communications
- Whales Don’t Ask for Directions: Here’s How They Always Know the Way (2025) – Whale Scientists
- Monitoring whale migration from space (2025) – WWF Arctic
- Whisperers of whales wander: A directional statistical investigation of whales’ migration influenced by geomagnetic, ocean current, and celestial cues (2025) – Journal for Nature Conservation
- Timing is everything: Drivers of interannual variability in blue whale migration (2020) – Scientific Reports
- Gray whales strand more often on days with increased levels of atmospheric radio-frequency noise (2020) – Current Biology
- Straight as an arrow: humpback whales swim constant course tracks during long-distance migration (2011) – Biology Letters
- Evidence from strandings for geomagnetic sensitivity in cetaceans (1986) – Journal of Experimental Biology
- Gray Whale Migration – NOAA Ocean Today
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 2 · Ecosystem engineers
Articles and studies
- Migrating baleen whales transport high-latitude nutrients to tropical and subtropical ecosystems (2025) – Nature Communications
- The Great Whale Conveyor Belt: Earth’s Largest Mammals Keep Oceans Thriving (2025) – Whale Scientists
- Opportunities, research gaps, and risks in allogenic ecosystem engineer mimicry (2025) – PubMed Central
- Whales and Carbon Sequestration: Can Whales Store Carbon? (2024) – NOAA Fisheries
- Whales are the engineers of their ecosystems (2021) – Whale Scientists
- Whales are Vital for Marine Ecosystem Health (2016) – International Whaling Commission
- Whales as marine ecosystem engineers (2014) – Frontiers in Ecology and the Environment
- Ecosystem Engineers – ScienceDirect Topics
- Blue Carbon – NOAA Office for Coastal Management
Lesson 3 · Diving
Articles and studies
- How To Study The Deepest Diving Whales? Use Chemistry! (2024) – Whale Scientists
- How did whales become the world’s deepest-diving mammals? (2021) – Whale Scientists
- Extreme diving in mammals: first estimates of behavioural aerobic dive limits in Cuvier’s beaked whales (2020) – Journal of Experimental Biology
- The advantages of diving deep: Fin whales quadruple their energy intake when targeting deep krill patches (2019) – Functional Ecology
- The CROW: Campus Research and Observational Writings, Volume Two (2017) – University of Washington Bothell
- Adaptations 6: Problems with pressure (2012) – The University of Western Australia
- Deadly diving? Physiological and behavioural management of decompression stress in diving mammals (2012) – Proceedings of the Royal Society B
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 4 · Culture
Articles and studies
- Animal culture: conservation in a changing world (2025) – Philosophical Transactions of the Royal Society B
- Culture and conservation in baleen whales (2025) – Philosophical Transactions of the Royal Society B
- Cultural Transmission, Evolution, and Revolution in Vocal Displays: Insights From Bird and Whale Song (2020) – Frontiers in Psychology
- Dolphin girl gang uses sponges to catch prey and create a new culture (2020) – Whale Scientists
- The interplay between social networks and culture: theoretically and among whales and dolphins (2013) – Philosophical Transactions of the Royal Society B
- Spontaneous human speech mimicry by a cetacean (2012) – Current Biology
Books
- The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Under Threat
Lesson 1 · Whaling
Articles and studies
- Historical Review of Whaling (2020) – Springer (book chapter)
- Too Much Is Never Enough: The Cautionary Tale of Soviet Illegal Whaling (2014) – Marine Fisheries Review
- A Review of Changes in the Use of Whale Resources over Time in Japan, with a Specific Example of the Hand-harpoon Fishery of Nago, Okinawa Prefecture (2013) – Semantic Scholar
- A taxonomy of world whaling: operations, eras, and data sources (2003) – NOAA
- Modern Eskimo whaling in the Alaskan Arctic (1988) – IEEE Xplore
- The Technology of Whaling in Australian Waters in the 19th Century (1983) – Semantic Scholar
- The Development of Indigenous Whaling: Prehistoric and Historic Contexts – ResearchGate
- Commercial whaling – International Whaling Commission
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 2 · Bycatch and entanglement
Articles and studies
- The State of World Fisheries and Aquaculture 2024: Fisheries and aquaculture employment (2024) – FAO
- How ‘dolphin safe’ is canned tuna, really? (2021) – National Geographic
- Bycatch in gillnet fisheries threatens Critically Endangered small cetaceans and other aquatic megafauna (2019) – Endangered Species Research
- Entanglement is a costly life-history stage in large whales (2017) – Ecology and Evolution
- Welfare Implications of Cetacean Bycatch and Entanglements (2017) – Marine Mammal Welfare (book chapter)
- The Painful Side of Trap and Fixed Net Fisheries: Chronic Entanglement of Large Whales (2012) – Journal of Marine Biology
- Lethal entanglement in baleen whales (2011) – Diseases of Aquatic Organisms
- Bycatch – International Whaling Commission
Lesson 3 · Noise pollution
Articles and studies
- Pilot whales shout louder to be heard over noise pollution (2026) – Journal of Experimental Biology
- Avoidance, confusion or solitude? Modelling how noise pollution affects whale migration (2024) – Movement Ecology
- Approaches to reduce noise from ships operating in important killer whale habitats (2019) – Marine Pollution Bulletin
- Evolution of Marine Noise Pollution Management (2015) – Aquatic Mammals
- Evidence that ship noise increases stress in right whales (2012) – Proceedings of the Royal Society B
- The impact of ocean noise pollution on marine biodiversity (2008) – Lindy Weilgart
Lesson 4 · Plastic and chemical pollution
Articles and studies
- Plastics – the fast Facts 2024 (2024) – Plastics Europe
- Chemical Pollution and Diseases of Marine Mammals (2023) – Marine Mammals (book chapter)
- 7 Solutions To Ocean Plastic Pollution (2023) – Oceanic Society
- Predicting global killer whale population collapse from PCB pollution (2018) – Science
- Evidence that the Great Pacific Garbage Patch is rapidly accumulating plastic (2018) – Scientific Reports
- Population growth is limited by nutritional impacts on pregnancy success in endangered Southern Resident killer whales (Orcinus orca) (2017) – PLoS ONE
- Exposure to a mixture of toxic chemicals: implications for the health of endangered southern resident killer whales (2016) – NOAA
- The impact of debris on marine life (2015) – Marine Pollution Bulletin
- As main meal for sperm whales: Plastics debris (2013) – Marine Pollution Bulletin
- Cetaceans and Marine Debris: The Great Unknown (2012) – Journal of Marine Biology
- Editorial: Plastic Pollution – An Ocean Emergency – ResearchGate
- Global plastics production – Our World in Data
- Facts – Seaspiracy
Lesson 5 · Climate change
Articles and studies
- Potential synergistic impacts of climate change and anthropogenic activity on Arctic marine mammals (2026) – Environmental Reviews
- The Ocean and Climate Change (2025) – NASA Science
- Climate change and cetacean health: impacts and future directions (2022) – PubMed
- Climate change-accelerated ocean biodiversity loss & associated planetary health impacts (2022) – ScienceDirect
- Rapid climate-driven circulation changes threaten conservation of endangered North Atlantic right whales (2019) – Oceanography
- Living Planet Index (2016) – WWF
- ‘The Blob’ invades Pacific, flummoxing climate experts (2015) – Science
- Living Blue Planet Report (2015) – WWF
- Arctic amplification dominated by temperature feedbacks in contemporary climate models (2014) – Nature Geoscience
- Processes and impacts of Arctic amplification: A research synthesis (2011) – Global and Planetary Change
- Loss of Arctic sea ice causing punctuated change in sightings of killer whales (Orcinus orca) over the past century (2009) – Ecological Applications
- Ocean acidification: the other CO2 problem (2009) – Annual Review of Marine Science
- Potential effects of climate change on marine mammals (2006) – Oceanography and Marine Biology: An Annual Review
- Long-term decline in krill stock and increase in salps within the Southern Ocean (2004) – Nature
- How climate change impacts the ocean – IFAW
- How Narwhals Find Food: Hunting Beneath Arctic Sea Ice – Narwhal.org
Conservation in Action
Lesson 1 · The humpback recovery
Articles and studies
- Changes in age-related sexual selection in a humpback whale population recovering from exploitation (2026) – Current Biology
- From Harpoons to Hope – The Return of Humpback Whales (2025) – Greenpeace Australia Pacific
- Why humpback whales are coming back (2025) – EarthSky
- Bellwethers of change: population modelling of North Pacific humpback whales from 2002 through 2021 reveals shift from recovery to climate response (2024) – Royal Society Open Science
- Humpback Whales Have Made a Remarkable Recovery, Giving Us Hope for the Planet (2020) – TIME
- Humpback whales make stunning comeback in southern Africa (2019) – National Geographic
- Post-whaling Recovery: The Beautiful Story of Humpbacks (2019) – Baleines en direct
- Assessing the recovery of an Antarctic predator from historical exploitation (2019) – Royal Society Open Science
- High pregnancy rates in humpback whales (Megaptera novaeangliae) around the Western Antarctic Peninsula, evidence of a rapidly growing population (2018) – Royal Society Open Science
- Humpback Whales: An Endangered Species Act Success Story? (2016) – NOAA Fisheries
- The whale that inspired Greenpeace (2013) – OUPblog – Frank Zelko
- Humpback whale – International Whaling Commission
- Humpback whale – Center for Biological Diversity
- The recovery of North Atlantic right whales, Eubalaena glacialis, has been constrained by human-caused mortality (2018) – Royal Society Open Science
Lesson 2 · The vaquita
Articles and studies
- Saving the Vaquita (2025) – Sea Shepherd
- Results of the 2025 Vaquita Monitoring Effort (2025) – Sea Shepherd
- Chapter 13 – Vaquita Phocoena sinus Norris and McFarland, 1958 (2025) – ScienceDirect (book chapter)
- Addressing Illegal Transnational Trade of Totoaba and Its Role in the Possible Extinction of the Vaquita (2023) – Journal of International Wildlife Law & Policy
- A field effort to capture critically endangered vaquitas Phocoena sinus for protection from entanglement in illegal gillnets (2019) – Endangered Species Research
- Conservation of the vaquita Phocoena sinus (2006) – Mammal Review
- What is killing the vaquita? – Porpoise Conservation Society
- Vaquita – NOAA Fisheries
Books
- Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Lesson 3 · Marine protected areas
Articles and studies
- A New Era for Ocean Protection (2026) – The Nature Conservancy
- What is the High Seas Treaty and Why is it Important? (2026) – United Nations University
- 10% of the ocean now protected. We have four years to triple it (2026) – Oceanographic
- Dominica creates the world’s first Sperm Whale Reserve, a win for conservation and climate (2025) – One Earth
- Dominica Establishes World’s First Sperm Whale Reserve, a Boost for Climate, Biodiversity and the Local Economy (2023) – National Geographic Society
- Extensive Use of Habitat-Damaging Fishing Gears Inside Habitat-Protecting Marine Protected Areas (2022) – Frontiers in Marine Science
- The Important Marine Mammal Area Network: A Tool for Systematic Spatial Planning in Response to the Marine Mammal Habitat Conservation Crisis (2022) – Frontiers in Marine Science
- The MPA Guide: A framework to achieve global goals for the ocean (2021) – Science
- Elevated trawling inside protected areas undermines conservation outcomes in a global fishing hot spot (2018) – Science
- Place-based approaches to marine mammal conservation (2016) – Aquatic Conservation
- Dynamic ocean management: Defining and conceptualizing real-time management of the ocean (2015) – Marine Policy
- Global conservation outcomes depend on marine protected areas with five key features (2014) – Nature
- The Importance of Marine Protected Areas (MPAs) – National Geographic Education
- MPAtlas – Marine Conservation Institute
- The MPA Guide – MPAtlas
Lesson 4 · Responsible whale watching
Articles and studies
- Your guide to the best, most responsible whale watching destinations (2026) – Travel Counsellors
- What is responsible whale watching? – IWC Whale Watching Handbook
- Importance of whale watching: Protecting marine life – IFAW
- Responsible whale watching and whale welfare – ResearchGate
- Keeping whales in captivity is wrong – Responsible Travel
- Wild vs. Captivity – Animal Welfare Institute
Lesson 5 · Citizen science
Articles and studies
- From citizen science to AI models: Advancing cetacean vocalization automatic detection through multi-annotator campaigns (2024) – Ecological Informatics
- The WhaleReport Alert System: Mitigating threats to whales with citizen science (2024) – Biological Conservation
- Population changes in a whale breeding ground revealed by citizen science noninvasive genetics (2022) – Global Ecology and Conservation
- Cidadãos, mídia social e ciência: a Ciência Cidadã como ferramenta de engajamento público na pesquisa com cetáceos – Maricato et al. (in Portuguese)
- Happywhale – Happywhale
- OceanWatchers – ORCA
- Citizen science – Whale Wise
- Whale Report Alert System (WRAS) – Ocean Wise
The Deep Dive
The depth records on the Deep Dive page.
- Mechanics, hydrodynamics and energetics of blue whale lunge feeding: efficiency dependence on krill density (2011) – Goldbogen et al., Journal of Experimental Biology 214: 131–146 · blue whale, 52–315 m
- Freediving – no limit (men) – Guinness World Records · Herbert Nitsch, 214 m, 2007
- Respiration and deep diving in the bottlenose porpoise (1969) – Ridgway, Scronce & Kanwisher, Science 166: 1651–1654 · bottlenose dolphin
- How deep can dolphins dive? – Dolphin Communication Project · bottlenose dolphin
- An insight into southern ocean orcas (2018) – World Cetacean Alliance · orca, 1,087 m
- Movements and dive behaviour of a toothfish-depredating killer and sperm whale – Towers et al., ICES Journal of Marine Science · orca
- Deep-diving foraging behaviour of sperm whales (Physeter macrocephalus) (2006) – Watwood et al., Journal of Animal Ecology 75: 814–825 · sperm whale
- How do deep-diving sea creatures withstand huge pressure changes? (2002) – Ponganis & Kooyman, Scientific American · sperm whale, ~2,000 m
- Using narwhals as oceanographic sampling platforms in the high Arctic (2007) – Laidre & Heide-Jørgensen, Oceanography 20(4) · narwhal
- Deep-diving by narwhals Monodon monoceros: differences in foraging behavior between wintering areas? (2003) – Laidre et al., Marine Ecology Progress Series 261: 269–281 · narwhal
- First long-term behavioral records from Cuvier’s beaked whales (Ziphius cavirostris) reveal record-breaking dives (2014) – Schorr et al., PLOS ONE 9(3): e92633 · 2,992 m
- Longest dive by a mammal – Guinness World Records · Cuvier’s beaked whale, 222 minutes (Quick et al. 2020, Journal of Experimental Biology)
Side by Side
The lengths and weights on the Side by Side page. Imperial figures were converted to metres and tonnes. The blue whale (32 m, 170 tonnes) and the male sperm whale (18 m) follow the figures in the course lessons and their sources.
- Blue whale – NOAA Fisheries · up to 110 ft, 330,000 lb
- Humpback whale – NOAA Fisheries · up to about 60 ft, about 40 tons
- North Atlantic right whale – NOAA Fisheries · up to 52 ft, 140,000 lb
- Gray whale – NOAA Fisheries · 42–49 ft, about 90,000 lb
- Bowhead whale – NOAA Fisheries · up to 62 ft, 200,000 lb
- Sperm whale – NOAA Fisheries · 40 ft (females) to 52 ft (males), up to 45 tons
- Killer whale – NOAA Fisheries · up to 32 ft, 11 tons
- Cuvier’s beaked whale – NOAA Fisheries · 15–23 ft, 4,000–6,800 lb
- Long-finned pilot whale – NOAA Fisheries · 19–25 ft, 2,900–5,000 lb
- Common bottlenose dolphin – NOAA Fisheries · 6–13 ft, 300–1,400 lb
- Harbor porpoise – NOAA Fisheries · 5–5.5 ft, 135–170 lb
- Vaquita – NOAA Fisheries · 4–5 ft, 65–120 lb
- Inia geoffrensis – Handbook of the Mammals of the World (via Plazi) · Amazon river dolphin, males 219–255 cm, up to 207 kg
- Narwhal – NAMMCO · males up to about 460 cm and 1,645 kg
- Narwhal – Norwegian Polar Institute · tusk up to 3 m
Images & sounds
Photographers, recordists and licences.
Sound recordings
- NOAA. Cornell University, Cornell Lab of Ornithology, K. Lisa Yang Center for Conservation Bioacoustics. 2023. Blue Whale. NOAA Sounds of the Ocean – used in “The blue whale”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Humpback Whale. NOAA Sounds of the Ocean – used in “The humpback whale”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. North Atlantic Right Whale 1. NOAA Sounds of the Ocean – used in “Right whales”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Bottlenose Dolphin. NOAA Sounds of the Ocean – used in “Dolphins”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Sperm Whale. NOAA Sounds of the Ocean – used in “Sperm whale”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Cuvier’s Beaked Whale. NOAA Sounds of the Ocean – used in “Beaked whales”
- NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Pilot Whale. NOAA Sounds of the Ocean – used in “Pilot whales”
- NOAA. Scottish Association for Marine Science, Denise Risch. 2023. Harbor Porpoise. NOAA Sounds of the Ocean – used in “Porpoises”
- NOAA. Alfred Wegener Institute, Ilse Van Opzeeland. 2023. Killer Whale. NOAA Sounds of the Ocean – used in “Orca”
Photographs
- Photo: Andriy Nekrasov – Plastic and chemical pollution
- Photo: Ethan Daniels – Pilot whales
- Photo: Francesco Ungaro – Underwater living
- Photo: Hernan Segui – Migration
- Photo: Ivan Stecko – Underwater living
- Photo: NOAA Fisheries – Porpoises
- Photo: NOAA Fisheries, Southwest Fisheries Science Center – Whaling
- Photo: NOAA Photo Library, CC BY 2.0 – Beaked whales
- Photo: Paula Olson, NOAA – The vaquita
- Photo: Roland Edler, CC BY 4.0 – Beaked whales
- Photo: Vicki Beaver, NOAA Fisheries – The bowhead whale
- Photo: marnock, Pexels – Porpoises
- Adobe Stock – Mammals that chose the ocean
- Shutterstock – 28 lessons
- Photo: Adam Ernster, Pexels – orca breaching, home page
Diagrams
- Diagrams and charts in the lessons were made for this course.