Module one

What is a Cetacean?

Lesson 1 · Mammals that chose the ocean

Articles and studies

  1. A diminutive new basilosaurid whale reveals the trajectory of the cetacean life histories during the Eocene (2023) – Communications Biology
  2. Basilosaurus, the “King of Lizards” (2020) – Whale Scientists
  3. Hippos and Whales: Unlikely Cousins (2017) – Royal Ontario Museum
  4. Science Word of the Day: Involucrum (2015) – National Geographic
  5. Pakicetus (2022) – EBSCO Research Starters
  6. From Land to Water: the Origin of Whales, Dolphins, and Porpoises (2009) – Evolution: Education and Outreach
  7. New Protocetid Whale from the Middle Eocene of Pakistan: Birth on Land, Precocial Development, and Sexual Dimorphism (2009) – PLoS ONE
  8. Whales originated from aquatic artiodactyls in the Eocene epoch of India (2007) – Nature
  9. The Origin of Whales and the Power of Independent Evidence (2000) – NCSE
  10. Phylogenetic relationships among cetartiodactyls based on insertions of short and long interspersed elements: Hippopotamuses are the closest extant relatives of whales (1999) – PNAS
  11. The evolution of whales – Understanding Evolution, UC Berkeley

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
  2. Spying on Whales (2018) – Nick Pyenson
  3. The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell

Lesson 2 · Built for water

Articles and studies

  1. A Whale of a Tail: Unraveling the evolutionary secrets of how whales and dolphins adapted their backbones for aquatic life (2024) – Harvard University
  2. The Biological Productivity of the Ocean (2012) – Nature Education
  3. Species review of Cuvier’s beaked whale, Ziphius cavirostris (2011) – International Whaling Commission
  4. 5.1 Properties of Water – Introduction to Oceanography
  5. Properties of Water – ResearchGate
  6. Whales – Smithsonian Ocean
  7. The loudest voice in the animal kingdom – BBC Earth
  8. Do whales and dolphins have hair? – Whale and Dolphin Conservation
  9. Mammal – ScienceDirect Topics

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
  2. The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
  3. Cetacean Societies: Field Studies of Dolphins and Whales (2000) – Mann, Connor, Tyack & Whitehead (eds.)

Lesson 3 · Two great lineages

Articles and studies

  1. Cetaceans Have Complex Brains for Complex Cognition (2007) – PLoS Biology
  2. Intro to Odontocetes & Mysticetes (2017) – Ocean Conservation Research
  3. Vaquita – NOAA Fisheries
  4. Intelligence, Cognition and Emotion in Cetaceans – The Whale Sanctuary Project
  5. Are Whales Sentient? The Scientific Evidence (2026) – Biology Insights

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
  2. Cetacean Societies: Field Studies of Dolphins and Whales (2000) – Mann, Connor, Tyack & Whitehead (eds.)

Lesson 4 · Underwater living

Articles and studies

  1. Nursing Behavior in Sperm Whales (Physeter macrocephalus) (2023) – Animal Behavior and Cognition
  2. Fight or flight: antipredator strategies of baleen whales (2008) – Mammal Review
  3. Cetacean sleep: An unusual form of mammalian sleep (2008) – Neuroscience and Biobehavioral Reviews
  4. Lactation in Whales and Dolphins: Evidence of Divergence Between Baleen- and Toothed-Species (1997) – Journal of Mammary Gland Biology and Neoplasia
  5. [TITLE MISSING] – PubMed Central
  6. Blue Whale Life Cycle: Birth, Growth, and Lifespan (2026) – Science Insights
  7. Intelligence, Cognition and Emotion in Cetaceans – The Whale Sanctuary Project

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
Module two

The Great Filter Feeders

Lesson 1 · The blue whale

Articles and studies

  1. Colossal ears? How baleen whales hear low-frequency sound (2025) – Hearing Research
  2. Blue whales (2020) – Current Biology
  3. Insight into the kinematics of blue whale surface foraging through drone observations and prey data (2020) – PeerJ
  4. Balaenoptera musculus (Blue Whale) (2018) – IUCN Red List
  5. A pulsed-air model of blue whale B call vocalizations (2017) – Scientific Reports
  6. Worldwide decline in tonal frequencies of blue whale songs (2009) – Endangered Species Research
  7. Studying How Cetaceans Use Sound to Explore Their Environment (1997) – Perspectives in Ethology
  8. Blue Whale – NOAA Fisheries
  9. Blue whale – International Whaling Commission

Lesson 2 · The humpback whale

Articles and studies

  1. Southern Ocean humpback whales are shifting to an earlier return migration (2025) – Scientific Reports
  2. Study finds humpback whales only ones capable of bubble-net feeding (2025) – University of Hawaiʻi
  3. Whales always had voices. Roger Payne helped the world hear them (2023) – Greenpeace
  4. Why Send Whale Song Into Space? (2017) – Literary Hub
  5. Humpback whales interfering when mammal-eating killer whales attack other species: Mobbing behavior and interspecific altruism? (2017) – Marine Mammal Science
  6. Cultural revolution in whale songs (2000) – Nature
  7. How do the ventral grooves of rorquals function? – Baleines en direct

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)
  2. The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Module three

Toothed Whales

Lesson 3 · Beaked whales

Articles and studies

  1. Navigating the unknown: assessing anthropogenic threats to beaked whales, family Ziphiidae (2024) – Royal Society Open Science
  2. Behavioural Ecology Traits of Elusive Deep-Diver Whales Unravel a Complex Social Structure Influenced by Female Philopatry and Defence Polygyny (2022) – Frontiers in Marine Science
  3. Fear of Killer Whales Drives Extreme Synchrony in Deep Diving Beaked Whales (2020) – Scientific Reports
  4. Diving behaviour of Cuvier’s beaked whales (Ziphius cavirostris) off Cape Hatteras, North Carolina (2019) – Royal Society Open Science
  5. Advances in research on the impacts of anti-submarine sonar on beaked whales (2019) – Proceedings of the Royal Society B
  6. Microplastic and macroplastic ingestion by a deep diving, oceanic cetacean: The True’s beaked whale Mesoplodon mirus (2015) – Environmental Pollution
  7. No mass strandings since sonar ban (2013) – Nature
  8. Beaked whales respond to simulated and actual navy sonar (2011) – PLoS ONE
  9. Correlating military sonar use with beaked whale mass strandings: what do the historical data show? (2009) – Aquatic Mammals
  10. Extreme diving of beaked whales (2006) – Journal of Experimental Biology
  11. Elements of beaked whale anatomy and diving physiology and some hypothetical causes of sonar-related stranding (2006) – Journal of Cetacean Research and Management
  12. Understanding the impacts of anthropogenic sound on beaked whales (2005) – Journal of Cetacean Research and Management

Lessons 5–6 · Porpoises & River dolphins

Articles and studies

  1. Estimating the abundance of the critically endangered Baltic Proper harbour porpoise (Phocoena phocoena) population using passive acoustic monitoring (2022) – Ecology and Evolution
  2. A case of playful interaction between Bolivian river dolphins with a Beni anaconda (2022) – JSTOR
  3. River Dolphins Spotted in Rare Playful Interaction With a Beni Anaconda (2022) – Smithsonian Magazine
  4. Polluted porpoises: Generational transfer of organic contaminants in harbour porpoises from the southern North Sea (2021) – Science of the Total Environment
  5. Interacting effects of vessel noise and shallow river depth elevate metabolic stress in Ganges river dolphins (2019) – Scientific Reports
  6. Porpoises the World Over: Diversity in Behavior and Ecology (2019) – Ethology and Behavioral Ecology of Odontocetes (book chapter)
  7. High field metabolic rates of wild harbour porpoises (2018) – Journal of Experimental Biology
  8. Morphological convergence in ‘river dolphin’ skulls (2017) – PeerJ
  9. Ultra-High Foraging Rates of Harbor Porpoises Make Them Vulnerable to Anthropogenic Disturbance (2016) – Current Biology
  10. The Neocortex of Indian River Dolphins (Genus Platanista): Comparative, Qualitative and Quantitative Analysis (2016) – Brain, Behavior and Evolution
  11. Do Porpoises Choose Their Associates? A New Method for Analyzing Social Relationships among Cetaceans (2011) – PLoS ONE
  12. Coexistence of Fisheries with River Dolphin Conservation (2010) – Conservation Biology
  13. Bye Baiji? (2007) – Current Biology
  14. River Dolphins Initiative – WWF
  15. Myths and Legends of the Amazon’s Pink River Dolphin – South America Vacations

Lesson 7 · Echolocation

Articles and studies

  1. For Dolphins, Echolocation May Be More Like ‘Touching’ Than ‘Seeing’ (2025) – Scientific American
  2. Adaptive echolocation behavior of bats and toothed whales in dynamic soundscapes (2023) – Journal of Experimental Biology
  3. Echolocating toothed whales use ultra-fast echo-kinetic responses to track evasive prey (2021) – eLife
  4. A bigger nose, a bigger bang: size matters for echolocating toothed whales (2018) – Aarhus University
  5. Clicking in Shallow Rivers: Short-Range Echolocation of Irrawaddy and Ganges River Dolphins in a Shallow, Acoustically Complex Habitat (2013) – PLoS ONE
  6. The Acoustic World of Harbor Porpoises – American Scientist
  7. Possible limitations of dolphin echolocation: a simulation study based on a cross-modal matching experiment – no link

Lesson 8 · Orca

Articles and studies

  1. Why these orcas are wearing salmon as hats (again) (2024) – National Geographic
  2. Ancestry testing of “Old Tom,” a killer whale central to mutualistic interactions with human whalers (2023) – Journal of Heredity
  3. These orcas control the waves to hunt. It’s spine-tingling to watch. (2023) – National Geographic
  4. Independent acoustic variation of the higher- and lower-frequency components of biphonic calls can facilitate call recognition and social affiliation in killer whales (2020) – PLoS ONE
  5. Genome-culture coevolution promotes rapid divergence of killer whale ecotypes (2016) – Nature Communications
  6. The Cooperation of Humans and Killer Whales (Orcinus orca): The Application of a Simple Fuzzy Rule-Based Model to a Historical System (2015) – Academia.edu
  7. Killer whale ecotypes: Is there a global model? (2013) – Biological Reviews
  8. Cultural traditions and the evolution of reproductive isolation: ecological speciation in killer whales? (2012) – Biological Journal of the Linnean Society
  9. Antarctic Peninsula killer whales (Orcinus orca) hunt seals and a penguin on floating ice (2012) – Marine Mammal Science
  10. Cross-cultural and cross-ecotype production of a killer whale ‘excitement’ call suggests universality (2011) – Naturwissenschaften
  11. Killer whales (Orcinus orca) feeding on schooling herring (Clupea harengus) using underwater tail-slaps: kinematic analyses of field observations (2000) – Journal of Experimental Biology
  12. Development of intentional stranding hunting techniques in killer whale (Orcinus orca) calves at Crozet Archipelago (1995) – Canadian Journal of Zoology
  13. Surface and underwater observations of cooperatively feeding killer whales in northern Norway (1993) – Canadian Journal of Zoology
  14. Intentional stranding apprenticeship and social play in killer whales (Orcinus orca) (1991) – Canadian Journal of Zoology
  15. Acoustic behaviour of resident killer whales (Orcinus orca) off Vancouver Island, British Columbia (1989) – Canadian Journal of Zoology

Books

  1. Endangered Orcas: The Story of the Southern Residents (2019) – Monika Wieland Shields
Module four

Lives in the Ocean

Lesson 2 · Ecosystem engineers

Articles and studies

  1. Migrating baleen whales transport high-latitude nutrients to tropical and subtropical ecosystems (2025) – Nature Communications
  2. The Great Whale Conveyor Belt: Earth’s Largest Mammals Keep Oceans Thriving (2025) – Whale Scientists
  3. Opportunities, research gaps, and risks in allogenic ecosystem engineer mimicry (2025) – PubMed Central
  4. Whales and Carbon Sequestration: Can Whales Store Carbon? (2024) – NOAA Fisheries
  5. Whales are the engineers of their ecosystems (2021) – Whale Scientists
  6. Whales are Vital for Marine Ecosystem Health (2016) – International Whaling Commission
  7. Whales as marine ecosystem engineers (2014) – Frontiers in Ecology and the Environment
  8. Ecosystem Engineers – ScienceDirect Topics
  9. Blue Carbon – NOAA Office for Coastal Management

Lesson 3 · Diving

Articles and studies

  1. How To Study The Deepest Diving Whales? Use Chemistry! (2024) – Whale Scientists
  2. How did whales become the world’s deepest-diving mammals? (2021) – Whale Scientists
  3. Extreme diving in mammals: first estimates of behavioural aerobic dive limits in Cuvier’s beaked whales (2020) – Journal of Experimental Biology
  4. The advantages of diving deep: Fin whales quadruple their energy intake when targeting deep krill patches (2019) – Functional Ecology
  5. The CROW: Campus Research and Observational Writings, Volume Two (2017) – University of Washington Bothell
  6. Adaptations 6: Problems with pressure (2012) – The University of Western Australia
  7. Deadly diving? Physiological and behavioural management of decompression stress in diving mammals (2012) – Proceedings of the Royal Society B

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)

Lesson 4 · Culture

Articles and studies

  1. Animal culture: conservation in a changing world (2025) – Philosophical Transactions of the Royal Society B
  2. Culture and conservation in baleen whales (2025) – Philosophical Transactions of the Royal Society B
  3. Cultural Transmission, Evolution, and Revolution in Vocal Displays: Insights From Bird and Whale Song (2020) – Frontiers in Psychology
  4. Dolphin girl gang uses sponges to catch prey and create a new culture (2020) – Whale Scientists
  5. The interplay between social networks and culture: theoretically and among whales and dolphins (2013) – Philosophical Transactions of the Royal Society B
  6. Spontaneous human speech mimicry by a cetacean (2012) – Current Biology

Books

  1. The Cultural Lives of Whales and Dolphins (2015) – Hal Whitehead & Luke Rendell
Module five

Under Threat

Lesson 1 · Whaling

Articles and studies

  1. Historical Review of Whaling (2020) – Springer (book chapter)
  2. Too Much Is Never Enough: The Cautionary Tale of Soviet Illegal Whaling (2014) – Marine Fisheries Review
  3. A Review of Changes in the Use of Whale Resources over Time in Japan, with a Specific Example of the Hand-harpoon Fishery of Nago, Okinawa Prefecture (2013) – Semantic Scholar
  4. A taxonomy of world whaling: operations, eras, and data sources (2003) – NOAA
  5. Modern Eskimo whaling in the Alaskan Arctic (1988) – IEEE Xplore
  6. The Technology of Whaling in Australian Waters in the 19th Century (1983) – Semantic Scholar
  7. The Development of Indigenous Whaling: Prehistoric and Historic Contexts – ResearchGate
  8. Commercial whaling – International Whaling Commission

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)

Lesson 2 · Bycatch and entanglement

Articles and studies

  1. The State of World Fisheries and Aquaculture 2024: Fisheries and aquaculture employment (2024) – FAO
  2. How ‘dolphin safe’ is canned tuna, really? (2021) – National Geographic
  3. Bycatch in gillnet fisheries threatens Critically Endangered small cetaceans and other aquatic megafauna (2019) – Endangered Species Research
  4. Entanglement is a costly life-history stage in large whales (2017) – Ecology and Evolution
  5. Welfare Implications of Cetacean Bycatch and Entanglements (2017) – Marine Mammal Welfare (book chapter)
  6. The Painful Side of Trap and Fixed Net Fisheries: Chronic Entanglement of Large Whales (2012) – Journal of Marine Biology
  7. Lethal entanglement in baleen whales (2011) – Diseases of Aquatic Organisms
  8. Bycatch – International Whaling Commission

Lesson 3 · Noise pollution

Articles and studies

  1. Pilot whales shout louder to be heard over noise pollution (2026) – Journal of Experimental Biology
  2. Avoidance, confusion or solitude? Modelling how noise pollution affects whale migration (2024) – Movement Ecology
  3. Approaches to reduce noise from ships operating in important killer whale habitats (2019) – Marine Pollution Bulletin
  4. Evolution of Marine Noise Pollution Management (2015) – Aquatic Mammals
  5. Evidence that ship noise increases stress in right whales (2012) – Proceedings of the Royal Society B
  6. The impact of ocean noise pollution on marine biodiversity (2008) – Lindy Weilgart

Lesson 5 · Climate change

Articles and studies

  1. Potential synergistic impacts of climate change and anthropogenic activity on Arctic marine mammals (2026) – Environmental Reviews
  2. The Ocean and Climate Change (2025) – NASA Science
  3. Climate change and cetacean health: impacts and future directions (2022) – PubMed
  4. Climate change-accelerated ocean biodiversity loss & associated planetary health impacts (2022) – ScienceDirect
  5. Rapid climate-driven circulation changes threaten conservation of endangered North Atlantic right whales (2019) – Oceanography
  6. Living Planet Index (2016) – WWF
  7. ‘The Blob’ invades Pacific, flummoxing climate experts (2015) – Science
  8. Living Blue Planet Report (2015) – WWF
  9. Arctic amplification dominated by temperature feedbacks in contemporary climate models (2014) – Nature Geoscience
  10. Processes and impacts of Arctic amplification: A research synthesis (2011) – Global and Planetary Change
  11. Loss of Arctic sea ice causing punctuated change in sightings of killer whales (Orcinus orca) over the past century (2009) – Ecological Applications
  12. Ocean acidification: the other CO2 problem (2009) – Annual Review of Marine Science
  13. Potential effects of climate change on marine mammals (2006) – Oceanography and Marine Biology: An Annual Review
  14. Long-term decline in krill stock and increase in salps within the Southern Ocean (2004) – Nature
  15. How climate change impacts the ocean – IFAW
  16. How Narwhals Find Food: Hunting Beneath Arctic Sea Ice – Narwhal.org
Module six

Conservation in Action

Lesson 1 · The humpback recovery

Articles and studies

  1. Changes in age-related sexual selection in a humpback whale population recovering from exploitation (2026) – Current Biology
  2. From Harpoons to Hope – The Return of Humpback Whales (2025) – Greenpeace Australia Pacific
  3. Why humpback whales are coming back (2025) – EarthSky
  4. Bellwethers of change: population modelling of North Pacific humpback whales from 2002 through 2021 reveals shift from recovery to climate response (2024) – Royal Society Open Science
  5. Humpback Whales Have Made a Remarkable Recovery, Giving Us Hope for the Planet (2020) – TIME
  6. Humpback whales make stunning comeback in southern Africa (2019) – National Geographic
  7. Post-whaling Recovery: The Beautiful Story of Humpbacks (2019) – Baleines en direct
  8. Assessing the recovery of an Antarctic predator from historical exploitation (2019) – Royal Society Open Science
  9. High pregnancy rates in humpback whales (Megaptera novaeangliae) around the Western Antarctic Peninsula, evidence of a rapidly growing population (2018) – Royal Society Open Science
  10. Humpback Whales: An Endangered Species Act Success Story? (2016) – NOAA Fisheries
  11. The whale that inspired Greenpeace (2013) – OUPblog – Frank Zelko
  12. Humpback whale – International Whaling Commission
  13. Humpback whale – Center for Biological Diversity
  14. The recovery of North Atlantic right whales, Eubalaena glacialis, has been constrained by human-caused mortality (2018) – Royal Society Open Science

Lesson 2 · The vaquita

Articles and studies

  1. Saving the Vaquita (2025) – Sea Shepherd
  2. Results of the 2025 Vaquita Monitoring Effort (2025) – Sea Shepherd
  3. Chapter 13 – Vaquita Phocoena sinus Norris and McFarland, 1958 (2025) – ScienceDirect (book chapter)
  4. Addressing Illegal Transnational Trade of Totoaba and Its Role in the Possible Extinction of the Vaquita (2023) – Journal of International Wildlife Law & Policy
  5. A field effort to capture critically endangered vaquitas Phocoena sinus for protection from entanglement in illegal gillnets (2019) – Endangered Species Research
  6. Conservation of the vaquita Phocoena sinus (2006) – Mammal Review
  7. What is killing the vaquita? – Porpoise Conservation Society
  8. Vaquita – NOAA Fisheries

Books

  1. Encyclopedia of Marine Mammals, 3rd edition (2018) – Würsig, Thewissen & Kovacs (eds.)

Lesson 3 · Marine protected areas

Articles and studies

  1. A New Era for Ocean Protection (2026) – The Nature Conservancy
  2. What is the High Seas Treaty and Why is it Important? (2026) – United Nations University
  3. 10% of the ocean now protected. We have four years to triple it (2026) – Oceanographic
  4. Dominica creates the world’s first Sperm Whale Reserve, a win for conservation and climate (2025) – One Earth
  5. Dominica Establishes World’s First Sperm Whale Reserve, a Boost for Climate, Biodiversity and the Local Economy (2023) – National Geographic Society
  6. Extensive Use of Habitat-Damaging Fishing Gears Inside Habitat-Protecting Marine Protected Areas (2022) – Frontiers in Marine Science
  7. The Important Marine Mammal Area Network: A Tool for Systematic Spatial Planning in Response to the Marine Mammal Habitat Conservation Crisis (2022) – Frontiers in Marine Science
  8. The MPA Guide: A framework to achieve global goals for the ocean (2021) – Science
  9. Elevated trawling inside protected areas undermines conservation outcomes in a global fishing hot spot (2018) – Science
  10. Place-based approaches to marine mammal conservation (2016) – Aquatic Conservation
  11. Dynamic ocean management: Defining and conceptualizing real-time management of the ocean (2015) – Marine Policy
  12. Global conservation outcomes depend on marine protected areas with five key features (2014) – Nature
  13. The Importance of Marine Protected Areas (MPAs) – National Geographic Education
  14. MPAtlas – Marine Conservation Institute
  15. The MPA Guide – MPAtlas

Lesson 4 · Responsible whale watching

Articles and studies

  1. Your guide to the best, most responsible whale watching destinations (2026) – Travel Counsellors
  2. What is responsible whale watching? – IWC Whale Watching Handbook
  3. Importance of whale watching: Protecting marine life – IFAW
  4. Responsible whale watching and whale welfare – ResearchGate
  5. Keeping whales in captivity is wrong – Responsible Travel
  6. Wild vs. Captivity – Animal Welfare Institute
Interactive

The Deep Dive

The depth records on the Deep Dive page.

  1. Mechanics, hydrodynamics and energetics of blue whale lunge feeding: efficiency dependence on krill density (2011) – Goldbogen et al., Journal of Experimental Biology 214: 131–146 · blue whale, 52–315 m
  2. Freediving – no limit (men) – Guinness World Records · Herbert Nitsch, 214 m, 2007
  3. Respiration and deep diving in the bottlenose porpoise (1969) – Ridgway, Scronce & Kanwisher, Science 166: 1651–1654 · bottlenose dolphin
  4. How deep can dolphins dive? – Dolphin Communication Project · bottlenose dolphin
  5. An insight into southern ocean orcas (2018) – World Cetacean Alliance · orca, 1,087 m
  6. Movements and dive behaviour of a toothfish-depredating killer and sperm whale – Towers et al., ICES Journal of Marine Science · orca
  7. Deep-diving foraging behaviour of sperm whales (Physeter macrocephalus) (2006) – Watwood et al., Journal of Animal Ecology 75: 814–825 · sperm whale
  8. How do deep-diving sea creatures withstand huge pressure changes? (2002) – Ponganis & Kooyman, Scientific American · sperm whale, ~2,000 m
  9. Using narwhals as oceanographic sampling platforms in the high Arctic (2007) – Laidre & Heide-Jørgensen, Oceanography 20(4) · narwhal
  10. Deep-diving by narwhals Monodon monoceros: differences in foraging behavior between wintering areas? (2003) – Laidre et al., Marine Ecology Progress Series 261: 269–281 · narwhal
  11. First long-term behavioral records from Cuvier’s beaked whales (Ziphius cavirostris) reveal record-breaking dives (2014) – Schorr et al., PLOS ONE 9(3): e92633 · 2,992 m
  12. Longest dive by a mammal – Guinness World Records · Cuvier’s beaked whale, 222 minutes (Quick et al. 2020, Journal of Experimental Biology)
Interactive

Side by Side

The lengths and weights on the Side by Side page. Imperial figures were converted to metres and tonnes. The blue whale (32 m, 170 tonnes) and the male sperm whale (18 m) follow the figures in the course lessons and their sources.

  1. Blue whale – NOAA Fisheries · up to 110 ft, 330,000 lb
  2. Humpback whale – NOAA Fisheries · up to about 60 ft, about 40 tons
  3. North Atlantic right whale – NOAA Fisheries · up to 52 ft, 140,000 lb
  4. Gray whale – NOAA Fisheries · 42–49 ft, about 90,000 lb
  5. Bowhead whale – NOAA Fisheries · up to 62 ft, 200,000 lb
  6. Sperm whale – NOAA Fisheries · 40 ft (females) to 52 ft (males), up to 45 tons
  7. Killer whale – NOAA Fisheries · up to 32 ft, 11 tons
  8. Cuvier’s beaked whale – NOAA Fisheries · 15–23 ft, 4,000–6,800 lb
  9. Long-finned pilot whale – NOAA Fisheries · 19–25 ft, 2,900–5,000 lb
  10. Common bottlenose dolphin – NOAA Fisheries · 6–13 ft, 300–1,400 lb
  11. Harbor porpoise – NOAA Fisheries · 5–5.5 ft, 135–170 lb
  12. Vaquita – NOAA Fisheries · 4–5 ft, 65–120 lb
  13. Inia geoffrensis – Handbook of the Mammals of the World (via Plazi) · Amazon river dolphin, males 219–255 cm, up to 207 kg
  14. Narwhal – NAMMCO · males up to about 460 cm and 1,645 kg
  15. Narwhal – Norwegian Polar Institute · tusk up to 3 m
Credits

Images & sounds

Photographers, recordists and licences.

Sound recordings

  1. NOAA. Cornell University, Cornell Lab of Ornithology, K. Lisa Yang Center for Conservation Bioacoustics. 2023. Blue Whale. NOAA Sounds of the Ocean – used in “The blue whale”
  2. NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Humpback Whale. NOAA Sounds of the Ocean – used in “The humpback whale”
  3. NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. North Atlantic Right Whale 1. NOAA Sounds of the Ocean – used in “Right whales”
  4. NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Bottlenose Dolphin. NOAA Sounds of the Ocean – used in “Dolphins”
  5. NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Sperm Whale. NOAA Sounds of the Ocean – used in “Sperm whale”
  6. NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Cuvier’s Beaked Whale. NOAA Sounds of the Ocean – used in “Beaked whales”
  7. NOAA. NOAA Fisheries, NEFSC, Passive Acoustics Branch. 2023. Pilot Whale. NOAA Sounds of the Ocean – used in “Pilot whales”
  8. NOAA. Scottish Association for Marine Science, Denise Risch. 2023. Harbor Porpoise. NOAA Sounds of the Ocean – used in “Porpoises”
  9. NOAA. Alfred Wegener Institute, Ilse Van Opzeeland. 2023. Killer Whale. NOAA Sounds of the Ocean – used in “Orca”

Photographs

  1. Photo: Andriy Nekrasov – Plastic and chemical pollution
  2. Photo: Ethan Daniels – Pilot whales
  3. Photo: Francesco Ungaro – Underwater living
  4. Photo: Hernan Segui – Migration
  5. Photo: Ivan Stecko – Underwater living
  6. Photo: NOAA Fisheries – Porpoises
  7. Photo: NOAA Fisheries, Southwest Fisheries Science Center – Whaling
  8. Photo: NOAA Photo Library, CC BY 2.0 – Beaked whales
  9. Photo: Paula Olson, NOAA – The vaquita
  10. Photo: Roland Edler, CC BY 4.0 – Beaked whales
  11. Photo: Vicki Beaver, NOAA Fisheries – The bowhead whale
  12. Photo: marnock, Pexels – Porpoises
  13. Adobe Stock – Mammals that chose the ocean
  14. Shutterstock – 28 lessons
  15. Photo: Adam Ernster, Pexels – orca breaching, home page

Diagrams

  1. Diagrams and charts in the lessons were made for this course.
Spotted a mistake? Science moves fast. If something in a lesson is out of date, I'd love to hear about it at [email protected].